At the time, the chemical hypothesis, based on analogy with Ephraim Racker's mechanism of substrate level phosphorylation linked to triose phosphate oxidation, seemed secure. He held the view that science progresses though open discussion, and abhorred the notion that ideas or information should be closeted away, hidden from "the competition". This view was captured in a nice remark he once made to me, that "The trouble with most scientists is not that they don't have good memories, but that they don't have good forgeteries." The field was evolving rapidly, and to those of us on the chemiosmotic side, the body of evidence favoring that point of view looked overwhelming. The Chemiosmotic Hypothesis.

In your own words, summarize the three hypothesis that were unified into Peter Mitchell’s Chemiosmotic Theory in 200-400 words.

metabolism, that led me to formulate the coupling hypothesis, summarised in Fig. I am grateful to Helen Mitchell for her kindness in supplying the picture of Peter Mitchell. The theory suggests essentially that most adenosine triphosphate (ATP) synthesis in respiring cells comes from the electrochemical gradient across the inner membranes of mitochondria by using the energy of NADH and FADH2 formed from the breaking down of energy-rich molecules such as glucose. It is interesting to look back and remember how sparse the clues were on which the hypothesis was based. In the wider context of his social and political views, Hayek was an early influence, and Peter would emphasize the role of the individual, and freedom of economic and political expression. Much of his thinking in the last 15 years was directed towards human and social problems, especially towards identifying mechanisms for conflict resolution.

[ But these must have seemed second-order problems to the main protagonists.

This hypothesis stated that a proton-motive force was responsible for driving the synthesis of ATP. Why did coupling seem so dependent on the maintenance of structure? The hypothesis found early favor among the photosynthetic community, perhaps because of the elegance of the early demonstrations from Jagendorf's lab, the explanation of amine uncoupling, the utility of the electrochromic "membrane voltmeters", perhaps also because of the more physico-chemical bent of the field. Discussions were a test bed for his latest ideas, and he relished the pursuit of odd angles and new perspectives.

Peter Mitchell proposed the _____ hypothesis that describes how _____ and _____ are coupled. Peter listed among his leisure interests (and here I quote from the International Who's Who), family life, home building, the creation of wealth and amenity, the restoration of buildings of architectural and historical interest, music, thinking, understanding, inventing, making, sailing. I first met Peter in 1962 when he visited Brian Chappell in Cambridge to talk mitochondriology. Without JavaScript you may encounter issues using the site.

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